481 lines
21 KiB
Markdown
481 lines
21 KiB
Markdown
# Tripartite Synapse — Pseudocode v11
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> Companion: `tripartite_synapse_v11_biology.md` explains the biology of every variable.
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> Changes from v10: each component now carries its own PARAMETERS and EXTERNAL INTERFACE
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> blocks; behaviors are organized into functional groups within each context; pool-filling
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> primitives return their increment (applied by the caller); the astrocyte no longer writes
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> PRE's drive (the spillover brake is PRE reading its own cleft).
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---
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## Functional groups (the uniform template, applied within each context)
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```
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RECEIVE take in resources + signals that arrived from outside
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BEHAVE the component's defining action
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EMIT send out — signals (messages) and resources (shipments) across the boundary
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SENSE deposit the fast trace that records the behavior
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RECOVER refill own private pools consumed by behaving
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EVALUATE judge the behavior — strength (needs dopamine) + endurance (interrupted success)
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DECAY let traces recede, closing their windows
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```
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Groups sit inside the DAY contexts (Option A): execution contexts (AP/bAP/CONTINUOUS) carry
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BEHAVE/EMIT/SENSE/endurance-EVALUATE; replenishment contexts (NOT_AP/NOT_bAP) carry
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RECEIVE/RECOVER/strength-EVALUATE/DECAY. Not every component uses every group.
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---
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## Conventions
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```
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SCOPE = {DAY, NIGHT} CONTEXT = {AP, NOT_AP, bAP, NOT_bAP, CONTINUOUS}
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DAY budget · fast_trace · possible_tag · endurance_need
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BRIDGE tag (POST: CANDIDATE→STABLE)
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NIGHT energy (not recoverable) · material (recoverable) · structure · budget_ceiling
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LOCALITY only local state + arrived signals; no component reads another's internal state.
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CLEFT MESSAGE CHANNELS (each component writes, others read)
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glutamate PRE → POST, ASTRO forward transmitter; ASTRO clears it
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astro_Dserine ASTRO → POST NMDA co-agonist gate
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retro_NO POST → PRE (+) "release reached a responsive target"
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retro_eCB POST → PRE (−) DSE: "over-driven, release less"
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SHIPMENT CHANNELS (source writes, target reads in its RECEIVE)
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soma_ship_dend SOMA→DEND soma_ship_axon SOMA→AXON
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dend_ship_post DEND→POST axon_ship_pre AXON→PRE
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```
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---
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## Primitives (return the increment; caller applies it)
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```
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sat(x, K) = x / (K + x)
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fill(pool, ceiling, rate, cost, budget) -> amount: // PRIVATE reserve
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amount = min(rate, ceiling - pool)·Δt
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budget -= amount·cost // intrinsic refill cost
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return amount
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refill(c from supply S) -> amount: // CONTESTED supply
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demand = c.budget_ceiling - c.budget
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factor = min(1, S / (Σ demand over components on S + ε))
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S -= demand·factor // consumes shared supply
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return demand·factor
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ship(from_budget, demand_sig, frac, cost) -> amount: // DIRECTED transfer
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amount = min(from_budget·frac, demand_sig) // demand_sig = propagated tag
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from_budget -= amount·(1 + cost)
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return amount // caller writes shipment channel
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```
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---
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## SHARED parameters (organism / physical / universal plasticity machinery)
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```
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// organism broadcasts (external; arrive as local levels)
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dopamine NE ACh
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// physical (external)
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glucose // vascular energy ceiling — root
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geometry // dendritic topology — bAP attenuation
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// universal plasticity gates (same mechanism in every component)
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elig dop_thr tag_thr tag_expiry // strength: eligibility → tag
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traj_thr endur_thr // endurance: interrupted-success → need
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// universal NIGHT
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decay_rate capacity_decay_rate recycle
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homeostatic_ceiling coherence_factor assembly_cost biogenesis_cost maint_cost
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// decay-time convention: each *= decay(τ) means relax toward 0 with time-constant τ
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```
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---
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---
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# DAY
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---
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## PRE
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```
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// PARAMETERS (pre)
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// K_release half-max Ca²⁺ for release release_cost gate to release
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// fusion_cost per unit NT_flux vatpase_cost per unit RRP refilled
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// spillover cleft level for autobrake brake autoreceptor factor
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//
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// EXTERNAL INTERFACE
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// EMIT glutamate → POST, ASTRO
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// RECEIVE astro_lactate[syn] ← ASTRO (primary fuel) ; axon_ship_pre ← AXON (fuel)
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// retro_NO, retro_eCB ← POST (backward messages)
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// pre_material ← AXON(NIGHT) ; pre_energy ← SOMA(NIGHT)
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// READ glutamate (own cleft, autobrake) ; dopamine (VTA, gates tag)
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// OWN pre_structure{slot_ceiling, VGCC_coupling, refill_ceiling} ; pre_budget_ceiling
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// EMERGENCY shockwave_lockdown ← ASTRO
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DAY | AP:
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// SENSE
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pre_fast_trace += spike_Ca(input_freq)
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// BEHAVE — release or fail
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if pre_budget < release_cost:
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suppress(NT_flux)
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// EVALUATE (endurance) — interrupted local success, confirmed by retro_NO
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if pre_fast_trace > traj_thr:
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pre_endurance_need += pre_fast_trace × (1 + retro_NO_local)
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exit
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drive = sat(pre_fast_trace, K_release) × (1 - retro_eCB_local)
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if RRP > 0:
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NT_flux = RRP × drive
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// EMIT
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glutamate += NT_flux·Δt
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RRP -= NT_flux·Δt; pre_budget -= NT_flux·fusion_cost
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if glutamate > spillover: drive *= brake // own-cleft autobrake
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DAY | NOT_AP:
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// RECEIVE
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retro_NO_local = retro_NO; retro_eCB_local = retro_eCB
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pre_budget += refill(pre from astro_lactate[syn] + axon_ship_pre)
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// RECOVER
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RRP += fill(RRP, pre_structure.slot_ceiling, pre_structure.refill_ceiling, vatpase_cost, pre_budget)
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// EVALUATE (strength)
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if pre_fast_trace > elig: pre_possible_tag += pre_fast_trace
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if dopamine > dop_thr and pre_possible_tag > tag_thr:
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pre_tag += dopamine × pre_possible_tag
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// DECAY
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pre_fast_trace *= decay(100ms); pre_possible_tag *= decay(s)
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pre_endurance_need *= decay(min); pre_tag *= decay(hr)
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dopamine *= decay(ms); retro_NO *= decay(s); retro_eCB *= decay(s)
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```
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---
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## POST
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```
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// PARAMETERS (post)
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// K_AMPA half-max glu for AMPA AMPA_Ca Ca fraction via AMPA
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// AMPA_cost NMDA_cost bAP_cost pka_cost traffic_cost req_cost
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// Mg_eject Vm to eject Mg block Dserine_thr NMDA co-agonist gate
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// Ca_STP fill-slots threshold Ca_TAG tagging threshold
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// eCB_thr Vm for endocannabinoid (DSE) drift baseline (STD relaxation)
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// NO_synth_cost eCB_synth_cost
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//
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// EXTERNAL INTERFACE
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// EMIT retro_NO (+), retro_eCB (−) → PRE
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// RECEIVE astro_lactate[syn] ← ASTRO ; dend_ship_post ← DEND
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// post_material ← DEND(NIGHT) ; post_energy ← SOMA(NIGHT)
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// READ glutamate ← PRE ; astro_Dserine ← ASTRO ; bAP (via dend_structure.bAP_fidelity)
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// dopamine (VTA, stabilizes tag)
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// OWN post_structure{slot_ceiling, spine_volume, reserve_ceiling} ; post_budget_ceiling
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// EMERGENCY shockwave_lockdown ← ASTRO
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DAY | NOT_bAP:
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// RECEIVE
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post_budget += refill(post from astro_lactate[syn] + dend_ship_post)
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// BEHAVE + SENSE — three Ca sources feed post_fast_trace
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a = sat(glutamate, K_AMPA) // SOURCE 1: AMPA
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AMPA_current = a × AMPA_surface; Vm += AMPA_current
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post_fast_trace += AMPA_Ca·AMPA_current; post_budget -= AMPA_cost
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if Vm > Mg_eject and astro_Dserine > Dserine_thr and glutamate > 0: // SOURCE 2: NMDA
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post_fast_trace += NMDA_Ca(glutamate)·rise_speed(); post_budget -= NMDA_cost
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// EMIT (+)
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retro_NO += NO_emit(post_fast_trace); post_budget -= NO_synth_cost
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// EMIT (−)
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if Vm > eCB_thr: retro_eCB += eCB_emit(Vm); post_budget -= eCB_synth_cost
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post_fast_trace *= decay(ms)
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// BEHAVE — STP fill slots (Ca-driven, private reserve); else STD drift (consequence)
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if post_fast_trace > Ca_STP:
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AMPA_surface = min(AMPA_surface + Ca_insert(post_fast_trace), post_structure.slot_ceiling)
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post_budget -= traffic_cost
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else:
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AMPA_surface = max(AMPA_surface - drift·Δt, baseline)
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// EVALUATE (endurance) — interrupted local success (own Ca climbing toward tag)
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if post_budget < req_cost and post_fast_trace > traj_thr and post_fast_trace_rising:
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post_endurance_need += post_fast_trace
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// EVALUATE (strength) — CANDIDATE then STABLE (dopamine)
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if post_fast_trace > Ca_TAG: post_possible_tag += post_fast_trace
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post_budget -= pka_cost
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if dopamine > dop_thr and post_possible_tag > tag_thr:
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post_tag += dopamine × post_possible_tag
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// DECAY
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post_possible_tag *= decay(min); post_endurance_need *= decay(min)
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post_tag *= decay(hr); dopamine *= decay(ms)
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DAY | bAP:
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// BEHAVE + SENSE — SOURCE 3: bAP amplifies an existing signal
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Vm += bAP_depol × dend_structure.bAP_fidelity; post_budget -= bAP_cost
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if post_possible_tag > Ca_TAG: post_fast_trace += bAP_Ca_boost()
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```
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---
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## DEND
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```
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// PARAMETERS (dend)
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// prop_cost branch_Ca_cost integrate_cost translate_cost
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//
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// EXTERNAL INTERFACE
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// EMIT bAP_local → POST ; branch_Vm → SOMA ; dend_ship_post → POST
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// RECEIVE astro_lactate[branch] ← ASTRO ; soma_ship_dend ← SOMA
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// dend_material, dend_energy ← SOMA(NIGHT)
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// READ SOMA.fired (bAP trigger) ; POST.Vm + spine spillover (to integrate)
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// dopamine ; ACh (commit-threshold context)
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// OWN dend_structure{bAP_fidelity(pos), translation_ceiling, transport_speed} ; dend_budget_ceiling
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DAY | bAP:
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// BEHAVE — propagate bAP (may fall short if depleted), integrate spines
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bAP_local, reached = propagate(SOMA.fired, dend_structure.bAP_fidelity, dend_budget, geometry)
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dend_budget -= prop_cost × reached
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// EVALUATE (endurance) — propagation cut short while branch strongly active (LOCAL)
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if reached < full and dend_fast_trace > traj_thr:
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dend_endurance_need += dend_fast_trace
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// SENSE
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dend_fast_trace += bAP_Ca(bAP_local) + spine_spillover(); dend_budget -= branch_Ca_cost
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// EMIT — propagated bAP to spines; integrated voltage to soma
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branch_Vm = integrate(POST.Vm, spines); dend_budget -= integrate_cost
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DAY | NOT_bAP:
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// RECEIVE
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dend_budget += refill(dend from astro_lactate[branch] + soma_ship_dend)
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// EMIT — ship budget downstream to spines (demand = post tag)
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dend_ship_post = ship(dend_budget, post_demand, post_ship_frac, ship_cost)
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// EVALUATE (strength)
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if dend_fast_trace > elig: dend_possible_tag += dend_fast_trace
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if dopamine > dop_thr and dend_possible_tag > tag_thr:
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dend_tag += dopamine × dend_possible_tag
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// BEHAVE — local translation if tagged (fills dend capacity faster)
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if dend_tag > tag_expiry and dend_budget > translate_cost: dend_budget -= translate_cost
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commit_threshold *= 1/(1 + ACh·gain)
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// DECAY
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dend_fast_trace *= decay(300ms); dend_possible_tag *= decay(s)
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dend_endurance_need *= decay(min); dend_tag *= decay(hr)
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```
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---
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## SOMA
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```
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// PARAMETERS (soma)
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// ap_cost nuclear_cost creb_cost mito_output (self-fuel rate)
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// inactivation ap_amp ap_contrib base_recovery
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// τ_Na τ_adapt τ_nuclear τ_align
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//
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// EXTERNAL INTERFACE
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// EMIT fired → AXON (propagate) + DEND (bAP) ; soma_ship_dend → DEND ; soma_ship_axon → AXON
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// RECEIVE self (mitochondria, ROOT) ; branch_Vm ← DEND
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// READ dopamine ; NE ; ACh
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// OWN soma_structure{baseline_threshold, AP_reliability, synthesis_ceiling} ; soma_budget_ceiling
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DAY | AP:
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threshold = soma_structure.baseline_threshold × (1 + soma_adaptation) × neuromod(NE, ACh)
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can_fire = soma_Na_inactivation < inactivation
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// BEHAVE — fire if able
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if branch_Vm > threshold and can_fire:
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if soma_budget < ap_cost:
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// EVALUATE (endurance) — fuel cut a firing train approaching CREB (LOCAL)
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if soma_fast_trace > traj_thr and soma_fast_trace_rising:
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soma_endurance_need += soma_fast_trace
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exit
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fired = True; soma_budget -= ap_cost // EMIT: fired → AXON, DEND
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// SENSE — three traces from one AP
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soma_Na_inactivation += ap_amp // → refractory (emergent)
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soma_adaptation += ap_contrib // → threshold rise
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soma_fast_trace += nuclear_Ca(); soma_budget -= nuclear_cost
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// EVALUATE (strength)
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if soma_fast_trace > elig: soma_possible_tag += soma_fast_trace
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if dopamine > dop_thr and soma_possible_tag > tag_thr:
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soma_tag += dopamine × soma_possible_tag
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soma_budget -= creb_cost
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DAY | NOT_AP:
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// BEHAVE — bottom-up refractory alignment (suprathreshold input during refractory)
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if branch_Vm > threshold and soma_Na_inactivation > inactivation:
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soma_refractory_alignment += (branch_Vm - threshold) × soma_Na_inactivation
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// RECOVER — recovery rate sped by alignment trace
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recovery = base_recovery × (1 + soma_refractory_alignment)
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soma_Na_inactivation *= decay(τ_Na / recovery)
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// RECEIVE — self-replenish (private root) + integrate input
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soma_budget += fill(soma_budget, soma_budget_ceiling, mito_output, 0, soma_budget)
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branch_Vm = integrate(DEND.branch_Vm, branches)
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// EMIT — ship downstream (demand = propagated tags)
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soma_ship_dend = ship(soma_budget, dend_demand, dend_ship_frac, ship_cost)
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soma_ship_axon = ship(soma_budget, axon_demand, axon_ship_frac, ship_cost)
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// DECAY
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soma_adaptation *= decay(τ_adapt); soma_fast_trace *= decay(τ_nuclear)
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soma_refractory_alignment *= decay(τ_align) // self-limiting
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soma_possible_tag *= decay(s); soma_endurance_need *= decay(min)
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soma_tag *= decay(hr); dopamine *= decay(ms)
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```
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---
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## AXON
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```
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// PARAMETERS (axon)
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// prop_cost budget_factor (depleted-propagation penalty)
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//
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// EXTERNAL INTERFACE
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// EMIT APs_delivered → PRE (propagation) ; axon_ship_pre → PRE
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// RECEIVE soma_ship_axon ← SOMA ; astro_lactate[shaft] ← ASTRO
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// axon_material, axon_energy ← SOMA(NIGHT)
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// READ SOMA.fired ; dopamine
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// OWN axon_structure{propagation, transport_ceiling, mito_density} ; axon_budget_ceiling
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DAY | AP:
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reliability = axon_structure.propagation × (1 - fail(axon_fast_trace))
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// BEHAVE — propagate (degraded if depleted)
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if axon_budget < prop_cost:
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reliability *= budget_factor
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// EVALUATE (endurance) — propagating a strong train when fuel failed (LOCAL)
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if axon_fast_trace > traj_thr: axon_endurance_need += axon_fast_trace
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delivered = fired × reliability; axon_budget -= prop_cost × delivered
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// EMIT — delivered APs reach boutons
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// SENSE
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axon_fast_trace += delivered; axon_fast_trace *= decay(s)
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DAY | NOT_AP:
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// RECEIVE
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axon_budget += refill(axon from soma_ship_axon + astro_lactate[shaft])
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// EMIT — ship budget to boutons (demand = pre tag)
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axon_ship_pre = ship(axon_budget, pre_demand, pre_ship_frac, ship_cost)
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// EVALUATE (strength)
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if axon_fast_trace > elig: axon_possible_tag += axon_fast_trace
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if dopamine > dop_thr and axon_possible_tag > tag_thr:
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axon_tag += dopamine × axon_possible_tag
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// DECAY
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axon_fast_trace *= decay(s); axon_possible_tag *= decay(s)
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axon_endurance_need *= decay(min); axon_tag *= decay(hr)
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```
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---
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## ASTRO
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```
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// PARAMETERS (astro)
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// K_Dserine half-max Ca for D-serine Ds_max Ds_frac Ds_cost
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// EAAT_cost per unit glutamate cleared lactate_cost per unit exported
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// spillover overflow threshold overload lockdown threshold
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//
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// EXTERNAL INTERFACE
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// EMIT astro_lactate[i] → pre/post/dend budgets ; astro_Dserine[i] → POST (gate)
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// RECEIVE glucose (ROOT) ; astro_material, astro_energy ← cell body (NIGHT)
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// READ glutamate ← PRE (clearance + spillover) ; dopamine
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// OWN astro_structure{perisynaptic_distance⁻¹, EAAT, Dserine_tonic, ECM} ; astro_budget_ceiling
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// EMERGENCY emits shockwave_lockdown on overload
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DAY | CONTINUOUS: // per astrosynapse i
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// RECEIVE — root production, capped by glucose
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astro_central_budget += glycolysis(glucose)·Δt
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// EMIT — demand-weighted lactate allocation across territory
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for each i: demand[i] = clearance_load[i] × astro_structure[i].delivery_eff
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factor = min(1, astro_central_budget / (Σ demand·lactate_cost + ε))
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for each i:
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astro_lactate[i] = demand[i] × factor; astro_central_budget -= astro_lactate[i]·lactate_cost
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// BEHAVE — clear glutamate; supply tonic D-serine
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glutamate[i] -= astro_structure[i].EAAT × glutamate[i]·Δt; astro_central_budget -= clearance·EAAT_cost
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astro_Dserine[i] += astro_structure[i].Dserine_tonic·Δt
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if glutamate[i] > spillover:
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// SENSE
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astro_fast_trace[i] += mGluR_Ca(); astro_fast_trace[i] *= decay(s)
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// BEHAVE + EMIT — D-serine pulse (demand-driven, budget-limited)
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want = sat(astro_fast_trace[i], K_Dserine) × Ds_max
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got = min(want, astro_central_budget × Ds_frac)
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astro_Dserine[i] += got; astro_central_budget -= got·Ds_cost
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// EVALUATE (endurance) — ran out of synthesis under high own demand (LOCAL)
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if got < want and astro_fast_trace[i] > traj_thr:
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astro_endurance_need[i] += (want - got)
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// EVALUATE (strength)
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if astro_fast_trace[i] > elig: astro_possible_tag[i] += astro_fast_trace[i]
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if dopamine > dop_thr and astro_possible_tag[i] > tag_thr:
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astro_tag[i] += dopamine × astro_possible_tag[i]
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// DECAY
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astro_possible_tag[i] *= decay(s); astro_endurance_need[i] *= decay(min)
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astro_tag[i] *= decay(hr)
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// EMERGENCY
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if astro_fast_trace[i] > overload: emit(shockwave_lockdown)
|
||
```
|
||
|
||
---
|
||
|
||
## Special — Shockwave Lockdown
|
||
|
||
```
|
||
DAY or NIGHT | OVERLOAD:
|
||
Vm = HYPERPOLARIZED; AMPA_surface = mass_internalize() → post reserve
|
||
axon_fast_trace += overdrive(); astro_central_budget -= emergency_cost
|
||
```
|
||
|
||
---
|
||
---
|
||
# NIGHT
|
||
System-wide consolidation (not a per-component DAY cycle). Build ceilings from DAY evidence.
|
||
|
||
```
|
||
NIGHT | 1 REPLENISH & DISTRIBUTE
|
||
// energy economy: astrocyte central → astrosynapses (astro_tag-weighted)
|
||
astro_central_{budget,energy,material} += overnight_*(glucose, …)·Δt
|
||
for each i with astro_tag[i] > tag_expiry: w = astro_tag[i]/Σastro_tag
|
||
astro_energy[i] += astro_central_energy·w; astro_material[i] += astro_central_material·w
|
||
// material economy: soma → branch/axon → spine/bouton
|
||
soma_budget += overnight_mito()·Δt; soma_energy += overnight_soma_energy()·Δt
|
||
soma_material += CREB_synth(soma_tag)·Δt // bottleneck
|
||
dend_material += soma_material·f_dend; axon_material += soma_material·f_axon; soma_material -= …
|
||
post_material += dend_material·f_spine; pre_material += axon_material·f_bouton
|
||
{pre,post,dend,axon}_energy += soma_energy·f[·]
|
||
{pre,post,dend,axon}_budget += astro_lactate_reserve·f[·]·Δt
|
||
|
||
NIGHT | 2 STRENGTH COMMITS (raise structure)
|
||
coherence = (pre_tag, post_tag, astro_tag all > tag_expiry) ? coherence_factor : 1
|
||
for each c with c_tag > tag_expiry:
|
||
Δ = min(slot_cost, c_material, c_energy·f)
|
||
c_structure += Δ × (coherence if c in {pre,post,astro} else 1)
|
||
c_material -= Δ; c_energy -= Δ·assembly_cost
|
||
if Δ < slot_cost: queue(c_strength_deficit → next NIGHT)
|
||
|
||
NIGHT | 2b ENDURANCE COMMITS (raise budget_ceiling) — no dopamine; competes with step 2
|
||
for each c with c_endurance_need > endur_thr:
|
||
Δ = min(cap_cost, c_material·f_cap, c_energy·f_cap)
|
||
c_budget_ceiling += Δ; c_material -= Δ; c_energy -= Δ·biogenesis_cost
|
||
if Δ < cap_cost: queue(c_endurance_deficit → next NIGHT)
|
||
|
||
NIGHT | 3 PASSIVE DECAY (both ceilings, by neglect)
|
||
maint = (total_material - consumed) × maint_frac / synapse_count
|
||
for each synapse:
|
||
{pre,post,dend,astro}_structure -= decay_rate·Δt
|
||
{pre,post,dend,astro}_budget_ceiling -= capacity_decay_rate·Δt
|
||
if maint ≥ maint_cost: structure += full_maint; budget_ceiling += full_cap_maint
|
||
else: structure += maint·frac; budget_ceiling += maint·cap_frac
|
||
for each synapse with net_change < 0: // material recovered, energy not
|
||
{pre,post,astro}_material += |net_change|·recycle·frac
|
||
|
||
NIGHT | 4 HOMEOSTATIC SCALING
|
||
if soma_tag > homeostatic_ceiling:
|
||
s = homeostatic_ceiling / soma_tag
|
||
for each synapse: post_structure.slot_ceiling *= s; pre_structure.slot_ceiling *= s
|
||
soma_material += Σ reduction·recycle
|
||
|
||
NIGHT | 5 CLEAR TRACES
|
||
all fast_trace, possible_tag, endurance_need = 0
|
||
soma_Na_inactivation = soma_adaptation = soma_refractory_alignment = 0
|
||
for each tag: if tag < tag_expiry: tag = 0 // else carry forward
|
||
// structure and budget_ceiling PERSIST
|
||
```
|
||
|
||
---
|
||
|
||
## One-view summary
|
||
|
||
```
|
||
DAY per context, per component: RECEIVE → BEHAVE/EMIT/SENSE → RECOVER → EVALUATE → DECAY
|
||
behavior runs within structure (strength) and budget_ceiling (endurance),
|
||
both filled by competition (refill = contested, fill = private, ship = directed)
|
||
fast_trace + dopamine coincidence → tag (strength evidence)
|
||
depletion + interrupted LOCAL success → endurance_need (endurance evidence)
|
||
NIGHT tag → structure ; endurance_need → budget_ceiling ; both draw one pool (compete)
|
||
unmaintained ceilings decay → freed material funds the rest
|
||
LOCAL every evaluation uses only own state + arrived signals; coupling is via channels.
|
||
```
|