diff --git a/elements/astrocyte/appunti/2026-06-06-tripartite_synapse_v2.md b/elements/astrocyte/appunti/2026-06-06-tripartite_synapse_v2.md
index c65721b..6287a65 100644
--- a/elements/astrocyte/appunti/2026-06-06-tripartite_synapse_v2.md
+++ b/elements/astrocyte/appunti/2026-06-06-tripartite_synapse_v2.md
@@ -723,7 +723,7 @@ Acetylcholine has two main sources: the basal forebrain nuclei (including the nu
 
 What's striking in the context of your model is that all three systems share the same architectural logic: a tiny, localized cell population broadcasts a global contextual signal that shifts the operational threshold of millions of synapses simultaneously — none of them carrying specific content, all of them modulating how content gets written.
 
-## Simple organisms
+### Simple organisms
 
 Excellent point. The hippocampal replay model is a vertebrate solution to a specific problem — how to consolidate many parallel experiences quickly without catastrophic interference. But the underlying molecular logic of the commit function is far more ancient and appears in organisms that have no hippocampus, no sleep architecture in the vertebrate sense, and sometimes no centralized nervous system at all.