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# Tripartite Synapse — Pseudocode v11
> Companion: `tripartite_synapse_v11_biology.md` explains the biology of every variable.
> Changes from v10: each component now carries its own PARAMETERS and EXTERNAL INTERFACE
> blocks; behaviors are organized into functional groups within each context; pool-filling
> primitives return their increment (applied by the caller); the astrocyte no longer writes
> PRE's drive (the spillover brake is PRE reading its own cleft).
---
## Functional groups (the uniform template, applied within each context)
```
RECEIVE take in resources + signals that arrived from outside
BEHAVE the component's defining action
EMIT send out — signals (messages) and resources (shipments) across the boundary
SENSE deposit the fast trace that records the behavior
RECOVER refill own private pools consumed by behaving
EVALUATE judge the behavior — strength (needs dopamine) + endurance (interrupted success)
DECAY let traces recede, closing their windows
```
Groups sit inside the DAY contexts (Option A): execution contexts (AP/bAP/CONTINUOUS) carry
BEHAVE/EMIT/SENSE/endurance-EVALUATE; replenishment contexts (NOT_AP/NOT_bAP) carry
RECEIVE/RECOVER/strength-EVALUATE/DECAY. Not every component uses every group.
---
## Conventions
```
SCOPE = {DAY, NIGHT} CONTEXT = {AP, NOT_AP, bAP, NOT_bAP, CONTINUOUS}
DAY budget · fast_trace · possible_tag · endurance_need
BRIDGE tag (POST: CANDIDATE→STABLE)
NIGHT energy (not recoverable) · material (recoverable) · structure · budget_ceiling
LOCALITY only local state + arrived signals; no component reads another's internal state.
CLEFT MESSAGE CHANNELS (each component writes, others read)
glutamate PRE → POST, ASTRO forward transmitter; ASTRO clears it
astro_Dserine ASTRO → POST NMDA co-agonist gate
retro_NO POST → PRE (+) "release reached a responsive target"
retro_eCB POST → PRE () DSE: "over-driven, release less"
SHIPMENT CHANNELS (source writes, target reads in its RECEIVE)
soma_ship_dend SOMA→DEND soma_ship_axon SOMA→AXON
dend_ship_post DEND→POST axon_ship_pre AXON→PRE
```
---
## Primitives (return the increment; caller applies it)
```
sat(x, K) = x / (K + x)
fill(pool, ceiling, rate, cost, budget) -> amount: // PRIVATE reserve
amount = min(rate, ceiling - pool)·Δt
budget -= amount·cost // intrinsic refill cost
return amount
refill(c from supply S) -> amount: // CONTESTED supply
demand = c.budget_ceiling - c.budget
factor = min(1, S / (Σ demand over components on S + ε))
S -= demand·factor // consumes shared supply
return demand·factor
ship(from_budget, demand_sig, frac, cost) -> amount: // DIRECTED transfer
amount = min(from_budget·frac, demand_sig) // demand_sig = propagated tag
from_budget -= amount·(1 + cost)
return amount // caller writes shipment channel
```
---
## SHARED parameters (organism / physical / universal plasticity machinery)
```
// organism broadcasts (external; arrive as local levels)
dopamine NE ACh
// physical (external)
glucose // vascular energy ceiling — root
geometry // dendritic topology — bAP attenuation
// universal plasticity gates (same mechanism in every component)
elig dop_thr tag_thr tag_expiry // strength: eligibility → tag
traj_thr endur_thr // endurance: interrupted-success → need
// universal NIGHT
decay_rate capacity_decay_rate recycle
homeostatic_ceiling coherence_factor assembly_cost biogenesis_cost maint_cost
// decay-time convention: each *= decay(τ) means relax toward 0 with time-constant τ
```
---
---
# DAY
---
## PRE
```
// PARAMETERS (pre)
// K_release half-max Ca²⁺ for release release_cost gate to release
// fusion_cost per unit NT_flux vatpase_cost per unit RRP refilled
// spillover cleft level for autobrake brake autoreceptor factor
//
// EXTERNAL INTERFACE
// EMIT glutamate → POST, ASTRO
// RECEIVE astro_lactate[syn] ← ASTRO (primary fuel) ; axon_ship_pre ← AXON (fuel)
// retro_NO, retro_eCB ← POST (backward messages)
// pre_material ← AXON(NIGHT) ; pre_energy ← SOMA(NIGHT)
// READ glutamate (own cleft, autobrake) ; dopamine (VTA, gates tag)
// OWN pre_structure{slot_ceiling, VGCC_coupling, refill_ceiling} ; pre_budget_ceiling
// EMERGENCY shockwave_lockdown ← ASTRO
DAY | AP:
// SENSE
pre_fast_trace += spike_Ca(input_freq)
// BEHAVE — release or fail
if pre_budget < release_cost:
suppress(NT_flux)
// EVALUATE (endurance) — interrupted local success, confirmed by retro_NO
if pre_fast_trace > traj_thr:
pre_endurance_need += pre_fast_trace × (1 + retro_NO_local)
exit
drive = sat(pre_fast_trace, K_release) × (1 - retro_eCB_local)
if RRP > 0:
NT_flux = RRP × drive
// EMIT
glutamate += NT_flux·Δt
RRP -= NT_flux·Δt; pre_budget -= NT_flux·fusion_cost
if glutamate > spillover: drive *= brake // own-cleft autobrake
DAY | NOT_AP:
// RECEIVE
retro_NO_local = retro_NO; retro_eCB_local = retro_eCB
pre_budget += refill(pre from astro_lactate[syn] + axon_ship_pre)
// RECOVER
RRP += fill(RRP, pre_structure.slot_ceiling, pre_structure.refill_ceiling, vatpase_cost, pre_budget)
// EVALUATE (strength)
if pre_fast_trace > elig: pre_possible_tag += pre_fast_trace
if dopamine > dop_thr and pre_possible_tag > tag_thr:
pre_tag += dopamine × pre_possible_tag
// DECAY
pre_fast_trace *= decay(100ms); pre_possible_tag *= decay(s)
pre_endurance_need *= decay(min); pre_tag *= decay(hr)
dopamine *= decay(ms); retro_NO *= decay(s); retro_eCB *= decay(s)
```
---
## POST
```
// PARAMETERS (post)
// K_AMPA half-max glu for AMPA AMPA_Ca Ca fraction via AMPA
// AMPA_cost NMDA_cost bAP_cost pka_cost traffic_cost req_cost
// Mg_eject Vm to eject Mg block Dserine_thr NMDA co-agonist gate
// Ca_STP fill-slots threshold Ca_TAG tagging threshold
// eCB_thr Vm for endocannabinoid (DSE) drift baseline (STD relaxation)
// NO_synth_cost eCB_synth_cost
//
// EXTERNAL INTERFACE
// EMIT retro_NO (+), retro_eCB () → PRE
// RECEIVE astro_lactate[syn] ← ASTRO ; dend_ship_post ← DEND
// post_material ← DEND(NIGHT) ; post_energy ← SOMA(NIGHT)
// READ glutamate ← PRE ; astro_Dserine ← ASTRO ; bAP (via dend_structure.bAP_fidelity)
// dopamine (VTA, stabilizes tag)
// OWN post_structure{slot_ceiling, spine_volume, reserve_ceiling} ; post_budget_ceiling
// EMERGENCY shockwave_lockdown ← ASTRO
DAY | NOT_bAP:
// RECEIVE
post_budget += refill(post from astro_lactate[syn] + dend_ship_post)
// BEHAVE + SENSE — three Ca sources feed post_fast_trace
a = sat(glutamate, K_AMPA) // SOURCE 1: AMPA
AMPA_current = a × AMPA_surface; Vm += AMPA_current
post_fast_trace += AMPA_Ca·AMPA_current; post_budget -= AMPA_cost
if Vm > Mg_eject and astro_Dserine > Dserine_thr and glutamate > 0: // SOURCE 2: NMDA
post_fast_trace += NMDA_Ca(glutamate)·rise_speed(); post_budget -= NMDA_cost
// EMIT (+)
retro_NO += NO_emit(post_fast_trace); post_budget -= NO_synth_cost
// EMIT ()
if Vm > eCB_thr: retro_eCB += eCB_emit(Vm); post_budget -= eCB_synth_cost
post_fast_trace *= decay(ms)
// BEHAVE — STP fill slots (Ca-driven, private reserve); else STD drift (consequence)
if post_fast_trace > Ca_STP:
AMPA_surface = min(AMPA_surface + Ca_insert(post_fast_trace), post_structure.slot_ceiling)
post_budget -= traffic_cost
else:
AMPA_surface = max(AMPA_surface - drift·Δt, baseline)
// EVALUATE (endurance) — interrupted local success (own Ca climbing toward tag)
if post_budget < req_cost and post_fast_trace > traj_thr and post_fast_trace_rising:
post_endurance_need += post_fast_trace
// EVALUATE (strength) — CANDIDATE then STABLE (dopamine)
if post_fast_trace > Ca_TAG: post_possible_tag += post_fast_trace
post_budget -= pka_cost
if dopamine > dop_thr and post_possible_tag > tag_thr:
post_tag += dopamine × post_possible_tag
// DECAY
post_possible_tag *= decay(min); post_endurance_need *= decay(min)
post_tag *= decay(hr); dopamine *= decay(ms)
DAY | bAP:
// BEHAVE + SENSE — SOURCE 3: bAP amplifies an existing signal
Vm += bAP_depol × dend_structure.bAP_fidelity; post_budget -= bAP_cost
if post_possible_tag > Ca_TAG: post_fast_trace += bAP_Ca_boost()
```
---
## DEND
```
// PARAMETERS (dend)
// prop_cost branch_Ca_cost integrate_cost translate_cost
//
// EXTERNAL INTERFACE
// EMIT bAP_local → POST ; branch_Vm → SOMA ; dend_ship_post → POST
// RECEIVE astro_lactate[branch] ← ASTRO ; soma_ship_dend ← SOMA
// dend_material, dend_energy ← SOMA(NIGHT)
// READ SOMA.fired (bAP trigger) ; POST.Vm + spine spillover (to integrate)
// dopamine ; ACh (commit-threshold context)
// OWN dend_structure{bAP_fidelity(pos), translation_ceiling, transport_speed} ; dend_budget_ceiling
DAY | bAP:
// BEHAVE — propagate bAP (may fall short if depleted), integrate spines
bAP_local, reached = propagate(SOMA.fired, dend_structure.bAP_fidelity, dend_budget, geometry)
dend_budget -= prop_cost × reached
// EVALUATE (endurance) — propagation cut short while branch strongly active (LOCAL)
if reached < full and dend_fast_trace > traj_thr:
dend_endurance_need += dend_fast_trace
// SENSE
dend_fast_trace += bAP_Ca(bAP_local) + spine_spillover(); dend_budget -= branch_Ca_cost
// EMIT — propagated bAP to spines; integrated voltage to soma
branch_Vm = integrate(POST.Vm, spines); dend_budget -= integrate_cost
DAY | NOT_bAP:
// RECEIVE
dend_budget += refill(dend from astro_lactate[branch] + soma_ship_dend)
// EMIT — ship budget downstream to spines (demand = post tag)
dend_ship_post = ship(dend_budget, post_demand, post_ship_frac, ship_cost)
// EVALUATE (strength)
if dend_fast_trace > elig: dend_possible_tag += dend_fast_trace
if dopamine > dop_thr and dend_possible_tag > tag_thr:
dend_tag += dopamine × dend_possible_tag
// BEHAVE — local translation if tagged (fills dend capacity faster)
if dend_tag > tag_expiry and dend_budget > translate_cost: dend_budget -= translate_cost
commit_threshold *= 1/(1 + ACh·gain)
// DECAY
dend_fast_trace *= decay(300ms); dend_possible_tag *= decay(s)
dend_endurance_need *= decay(min); dend_tag *= decay(hr)
```
---
## SOMA
```
// PARAMETERS (soma)
// ap_cost nuclear_cost creb_cost mito_output (self-fuel rate)
// inactivation ap_amp ap_contrib base_recovery
// τ_Na τ_adapt τ_nuclear τ_align
//
// EXTERNAL INTERFACE
// EMIT fired → AXON (propagate) + DEND (bAP) ; soma_ship_dend → DEND ; soma_ship_axon → AXON
// RECEIVE self (mitochondria, ROOT) ; branch_Vm ← DEND
// READ dopamine ; NE ; ACh
// OWN soma_structure{baseline_threshold, AP_reliability, synthesis_ceiling} ; soma_budget_ceiling
DAY | AP:
threshold = soma_structure.baseline_threshold × (1 + soma_adaptation) × neuromod(NE, ACh)
can_fire = soma_Na_inactivation < inactivation
// BEHAVE — fire if able
if branch_Vm > threshold and can_fire:
if soma_budget < ap_cost:
// EVALUATE (endurance) — fuel cut a firing train approaching CREB (LOCAL)
if soma_fast_trace > traj_thr and soma_fast_trace_rising:
soma_endurance_need += soma_fast_trace
exit
fired = True; soma_budget -= ap_cost // EMIT: fired → AXON, DEND
// SENSE — three traces from one AP
soma_Na_inactivation += ap_amp // → refractory (emergent)
soma_adaptation += ap_contrib // → threshold rise
soma_fast_trace += nuclear_Ca(); soma_budget -= nuclear_cost
// EVALUATE (strength)
if soma_fast_trace > elig: soma_possible_tag += soma_fast_trace
if dopamine > dop_thr and soma_possible_tag > tag_thr:
soma_tag += dopamine × soma_possible_tag
soma_budget -= creb_cost
DAY | NOT_AP:
// BEHAVE — bottom-up refractory alignment (suprathreshold input during refractory)
if branch_Vm > threshold and soma_Na_inactivation > inactivation:
soma_refractory_alignment += (branch_Vm - threshold) × soma_Na_inactivation
// RECOVER — recovery rate sped by alignment trace
recovery = base_recovery × (1 + soma_refractory_alignment)
soma_Na_inactivation *= decay(τ_Na / recovery)
// RECEIVE — self-replenish (private root) + integrate input
soma_budget += fill(soma_budget, soma_budget_ceiling, mito_output, 0, soma_budget)
branch_Vm = integrate(DEND.branch_Vm, branches)
// EMIT — ship downstream (demand = propagated tags)
soma_ship_dend = ship(soma_budget, dend_demand, dend_ship_frac, ship_cost)
soma_ship_axon = ship(soma_budget, axon_demand, axon_ship_frac, ship_cost)
// DECAY
soma_adaptation *= decay(τ_adapt); soma_fast_trace *= decay(τ_nuclear)
soma_refractory_alignment *= decay(τ_align) // self-limiting
soma_possible_tag *= decay(s); soma_endurance_need *= decay(min)
soma_tag *= decay(hr); dopamine *= decay(ms)
```
---
## AXON
```
// PARAMETERS (axon)
// prop_cost budget_factor (depleted-propagation penalty)
//
// EXTERNAL INTERFACE
// EMIT APs_delivered → PRE (propagation) ; axon_ship_pre → PRE
// RECEIVE soma_ship_axon ← SOMA ; astro_lactate[shaft] ← ASTRO
// axon_material, axon_energy ← SOMA(NIGHT)
// READ SOMA.fired ; dopamine
// OWN axon_structure{propagation, transport_ceiling, mito_density} ; axon_budget_ceiling
DAY | AP:
reliability = axon_structure.propagation × (1 - fail(axon_fast_trace))
// BEHAVE — propagate (degraded if depleted)
if axon_budget < prop_cost:
reliability *= budget_factor
// EVALUATE (endurance) — propagating a strong train when fuel failed (LOCAL)
if axon_fast_trace > traj_thr: axon_endurance_need += axon_fast_trace
delivered = fired × reliability; axon_budget -= prop_cost × delivered
// EMIT — delivered APs reach boutons
// SENSE
axon_fast_trace += delivered; axon_fast_trace *= decay(s)
DAY | NOT_AP:
// RECEIVE
axon_budget += refill(axon from soma_ship_axon + astro_lactate[shaft])
// EMIT — ship budget to boutons (demand = pre tag)
axon_ship_pre = ship(axon_budget, pre_demand, pre_ship_frac, ship_cost)
// EVALUATE (strength)
if axon_fast_trace > elig: axon_possible_tag += axon_fast_trace
if dopamine > dop_thr and axon_possible_tag > tag_thr:
axon_tag += dopamine × axon_possible_tag
// DECAY
axon_fast_trace *= decay(s); axon_possible_tag *= decay(s)
axon_endurance_need *= decay(min); axon_tag *= decay(hr)
```
---
## ASTRO
```
// PARAMETERS (astro)
// K_Dserine half-max Ca for D-serine Ds_max Ds_frac Ds_cost
// EAAT_cost per unit glutamate cleared lactate_cost per unit exported
// spillover overflow threshold overload lockdown threshold
//
// EXTERNAL INTERFACE
// EMIT astro_lactate[i] → pre/post/dend budgets ; astro_Dserine[i] → POST (gate)
// RECEIVE glucose (ROOT) ; astro_material, astro_energy ← cell body (NIGHT)
// READ glutamate ← PRE (clearance + spillover) ; dopamine
// OWN astro_structure{perisynaptic_distance⁻¹, EAAT, Dserine_tonic, ECM} ; astro_budget_ceiling
// EMERGENCY emits shockwave_lockdown on overload
DAY | CONTINUOUS: // per astrosynapse i
// RECEIVE — root production, capped by glucose
astro_central_budget += glycolysis(glucose)·Δt
// EMIT — demand-weighted lactate allocation across territory
for each i: demand[i] = clearance_load[i] × astro_structure[i].delivery_eff
factor = min(1, astro_central_budget / (Σ demand·lactate_cost + ε))
for each i:
astro_lactate[i] = demand[i] × factor; astro_central_budget -= astro_lactate[i]·lactate_cost
// BEHAVE — clear glutamate; supply tonic D-serine
glutamate[i] -= astro_structure[i].EAAT × glutamate[i]·Δt; astro_central_budget -= clearance·EAAT_cost
astro_Dserine[i] += astro_structure[i].Dserine_tonic·Δt
if glutamate[i] > spillover:
// SENSE
astro_fast_trace[i] += mGluR_Ca(); astro_fast_trace[i] *= decay(s)
// BEHAVE + EMIT — D-serine pulse (demand-driven, budget-limited)
want = sat(astro_fast_trace[i], K_Dserine) × Ds_max
got = min(want, astro_central_budget × Ds_frac)
astro_Dserine[i] += got; astro_central_budget -= got·Ds_cost
// EVALUATE (endurance) — ran out of synthesis under high own demand (LOCAL)
if got < want and astro_fast_trace[i] > traj_thr:
astro_endurance_need[i] += (want - got)
// EVALUATE (strength)
if astro_fast_trace[i] > elig: astro_possible_tag[i] += astro_fast_trace[i]
if dopamine > dop_thr and astro_possible_tag[i] > tag_thr:
astro_tag[i] += dopamine × astro_possible_tag[i]
// DECAY
astro_possible_tag[i] *= decay(s); astro_endurance_need[i] *= decay(min)
astro_tag[i] *= decay(hr)
// EMERGENCY
if astro_fast_trace[i] > overload: emit(shockwave_lockdown)
```
---
## Special — Shockwave Lockdown
```
DAY or NIGHT | OVERLOAD:
Vm = HYPERPOLARIZED; AMPA_surface = mass_internalize() → post reserve
axon_fast_trace += overdrive(); astro_central_budget -= emergency_cost
```
---
---
# NIGHT
System-wide consolidation (not a per-component DAY cycle). Build ceilings from DAY evidence.
```
NIGHT | 1 REPLENISH & DISTRIBUTE
// energy economy: astrocyte central → astrosynapses (astro_tag-weighted)
astro_central_{budget,energy,material} += overnight_*(glucose, …)·Δt
for each i with astro_tag[i] > tag_expiry: w = astro_tag[i]/Σastro_tag
astro_energy[i] += astro_central_energy·w; astro_material[i] += astro_central_material·w
// material economy: soma → branch/axon → spine/bouton
soma_budget += overnight_mito()·Δt; soma_energy += overnight_soma_energy()·Δt
soma_material += CREB_synth(soma_tag)·Δt // bottleneck
dend_material += soma_material·f_dend; axon_material += soma_material·f_axon; soma_material -= …
post_material += dend_material·f_spine; pre_material += axon_material·f_bouton
{pre,post,dend,axon}_energy += soma_energy·f[·]
{pre,post,dend,axon}_budget += astro_lactate_reserve·f[·]·Δt
NIGHT | 2 STRENGTH COMMITS (raise structure)
coherence = (pre_tag, post_tag, astro_tag all > tag_expiry) ? coherence_factor : 1
for each c with c_tag > tag_expiry:
Δ = min(slot_cost, c_material, c_energy·f)
c_structure += Δ × (coherence if c in {pre,post,astro} else 1)
c_material -= Δ; c_energy -= Δ·assembly_cost
if Δ < slot_cost: queue(c_strength_deficit → next NIGHT)
NIGHT | 2b ENDURANCE COMMITS (raise budget_ceiling) — no dopamine; competes with step 2
for each c with c_endurance_need > endur_thr:
Δ = min(cap_cost, c_material·f_cap, c_energy·f_cap)
c_budget_ceiling += Δ; c_material -= Δ; c_energy -= Δ·biogenesis_cost
if Δ < cap_cost: queue(c_endurance_deficit → next NIGHT)
NIGHT | 3 PASSIVE DECAY (both ceilings, by neglect)
maint = (total_material - consumed) × maint_frac / synapse_count
for each synapse:
{pre,post,dend,astro}_structure -= decay_rate·Δt
{pre,post,dend,astro}_budget_ceiling -= capacity_decay_rate·Δt
if maint ≥ maint_cost: structure += full_maint; budget_ceiling += full_cap_maint
else: structure += maint·frac; budget_ceiling += maint·cap_frac
for each synapse with net_change < 0: // material recovered, energy not
{pre,post,astro}_material += |net_change|·recycle·frac
NIGHT | 4 HOMEOSTATIC SCALING
if soma_tag > homeostatic_ceiling:
s = homeostatic_ceiling / soma_tag
for each synapse: post_structure.slot_ceiling *= s; pre_structure.slot_ceiling *= s
soma_material += Σ reduction·recycle
NIGHT | 5 CLEAR TRACES
all fast_trace, possible_tag, endurance_need = 0
soma_Na_inactivation = soma_adaptation = soma_refractory_alignment = 0
for each tag: if tag < tag_expiry: tag = 0 // else carry forward
// structure and budget_ceiling PERSIST
```
---
## One-view summary
```
DAY per context, per component: RECEIVE → BEHAVE/EMIT/SENSE → RECOVER → EVALUATE → DECAY
behavior runs within structure (strength) and budget_ceiling (endurance),
both filled by competition (refill = contested, fill = private, ship = directed)
fast_trace + dopamine coincidence → tag (strength evidence)
depletion + interrupted LOCAL success → endurance_need (endurance evidence)
NIGHT tag → structure ; endurance_need → budget_ceiling ; both draw one pool (compete)
unmaintained ceilings decay → freed material funds the rest
LOCAL every evaluation uses only own state + arrived signals; coupling is via channels.
```